At Camp Half-Blood, Percy Jackson (Logan Lerman) meets his half-brother Tyson (Douglas Smith), who is a cyclops. The camp is later attacked byLuke Castellan (Jake Abel), who announces his plans to destroy Mount Olympus. Percy's mentor Chiron (Anthony Head) discovers that Luke has poisoned the magic tree responsible for the barrier that protects Camp Half-Blood, which Percy learns was created out of Thalia Grace (Paloma Kwiatkowski), daughter of Zeus, who was killed by a cyclops. Annabeth Chase (Alexandra Daddario) finds out that the Golden Fleece could restore the tree, and the camp's director, Dionysus (Stanley Tucci), sends Clarisse La Rue (Leven Rambin), daughter of Ares and Percy's rival, to find it. Percy then sets off with Annabeth, Tyson and Grover Underwood (Brandon T. Jackson) to locate the Golden Fleece on his own. Before they leave, the Oracle (Shohreh Aghdashloo) prophesies that either a half-blood child of one of the three prime gods will fight Luke, and the half-blood will have on their hands the chance to either save Mount Olympus or destroy it. As Percy is the only known half-blood of the three prime gods (Tyson is not considered a half-blood because he is half cyclops), he realizes the prophecy must refer to him.
The Graeae (Missi Pyle, Yvette Nicole Brown and Mary Birdsong) give the group the coordinates to the island and leave them in Washington, D.C., where Grover is captured by Luke's men, who need him to find the Fleece as satyrs are naturally drawn to it. Grover fears the cyclops Polyphemus(Robert Maillet), the creature that guards the Golden Fleece and uses it to lure in satyrs to eat. Percy, Annabeth and Tyson then meet Luke's fatherHermes (Nathan Fillion), who tells them that Luke is in a ocean liner in the Atlantic Ocean called the Andromeda; he asks Percy to apologize on his behalf for being a bad father to Luke. Equipped with gifts of tape that makes things disappear and a thermos of wind from Hermes, Percy, Annabeth and Tyson take a Hippocampus to the Andromeda and end up being captured by one of Luke's soldiers, the Manticore (Daniel Cudmore), but escape using the magic artifacts. The trio eventually reaches the Sea of Monsters and is swallowed by Charybdis, meeting Clarisse in its stomach. She was given an old Civil War Confederate Ironclad from her father to use on her quest, run by a crew of Confederate Zombie Soldiers. They join forces to escape and reach Circeland, an abandoned amusement park above Polyphemus' lair, where they rescue Grover and retrieve the Golden Fleece before being confronted by Luke, who reveals his plans to use the Golden Fleece to awaken Kronos (Robert Knepper).
Tyson sacrifices himself to protect Percy from a crossbow bolt fired from Luke, who subsequently steals the Golden Fleece and awakens Kronos. Tyson then returns, having been revived in contact with water, and rescues the others. In the ensuing fight, Luke and Grover are swallowed by Kronos before Percy realizes his magic sword, Riptide, is prophesied to be Kronos' only weakness. Percy's sword swing send Kronos's body back, piece by piece, to the golden coffin which held his remains, and Luke becomes trapped in the hungry Polyphemus' lair. The Manticore mortally wounds Annabeth before being killed by Grover and Clarisse, and Percy uses the Golden Fleece to revive her.
Returning to Camp Half-Blood, the group uses the Golden Fleece to restore the tree, and are surprised when it revives Thalia and restores her body. While the others celebrate, Percy realizes that perhaps Mount Olympus' fate might rest on Thalia's hands, not his.
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